Last update: 23/11/20
Publications
[86] Fujimoto, Y. & Ohtsuki, H. "Who is a Leader in the Leading Eight? Indirect Reciprocity under Private Assessment"
PRX Life, (2024) 2, 023009, doi:10.1103/PRXLife.2.023009

[85] Parvinen, K., Ohtsuki, H. & Wakano, J.Y. "Evolution of dispersal under spatio-temporal heterogeneity"
Journal of Theoretical Biology, (2023) 574, 111612, doi:10.1016/j.jtbi.2023.111612

[84] Fujimoto, Y. & Ohtsuki, H. "Evolutionary stability of cooperation in indirect reciprocity under noisy and private assessment"
Proceedings of the National Academy of Sciences of the United States of America, (2023) 120(20), e2300544120, doi:10.1073/pnas.2300544120

[83] Sekiguchi, T. & Ohtsuki, H. "Aggregation of correlated judgments on multiple interconnected issues"
Group Decision and Negotiation, (2023) 32, 233-256, doi:10.1007/s10726-022-09806-w

[82] Kristensen, N.P., Ohtsuki, H. & Chisholm, R.A. "Ancestral social environments plus nonlinear benefits can explain cooperation in human societies"
Scientific Reports, (2022) 12, 20252, doi:10.1038/s41598-022-24590-y

[81] Fujimoto, Y. & Ohtsuki, H. "Reputation structure in indirect reciprocity under noisy and private assessment"
Scientific Reports, (2022) 12, 10500, doi:10.1038/s41598-022-14171-4

[80] Takeuchi, Y., Ohtsuki, H. & Innan, H. "A non-zero-sum neutrality test for the tropical rain forest community using long-term between-census data"
Ecology and Evolution, (2022) 12(1), e8462, doi:10.1002/ece3.8462

[79] Ohtsuki, H. & Ujiyama, S. "Impact of social dominance on the evolution of individual learning"
Journal of Theoretical Biology, (2022) 535, 110986, doi:10.1016/j.jtbi.2021.110986

[78] Mullon, C., Wakano, J.Y. & Ohtsuki, H. "Coevolutionary dynamics of genetic traits and their long-term extended effects under non-random interactions"
Journal of Theoretical Biology, (2021) 525, 110750, doi:10.1016/j.jtbi.2021.110750

[77] Konishi, S., Kariya, F., Hamasaki, K., Takayasu, L. & Ohtsuki, H. "Fecundability and Sterility by Age: Estimates Using Time to Pregnancy Data of Japanese Couples Trying to Conceive Their First Child with and without Fertility Treatment"
International Journal of Environmental Research and Public Health, (2021) 18(10), 5486, doi:10.3390/ijerph18105486

[76] Ohtsuki, H., Rueffler, C., Wakano, J.Y., Parvinen, K. & Lehmann, L. "The components of directional and disruptive selection in heterogeneous group-structured populations."
Journal of Theoretical Biology, (2020) 507, 110449, doi:10.1016/j.jtbi.2020.110449

[75] Nagasawa, M., Mogi, K., Ohtsuki, H. & Kikusui, T. "Familiarity with humans affect dogsf tendencies to follow human majority groups."
Scientific Reports, (2020) 10, 7119, doi:10.1038/s41598-020-64058-5

[74] Parvinen, K., Ohtsuki, H. & Wakano, J.Y. "Evolution of dispersal in a spatially heterogeneous population with finite patch sizes."
Proceedings of the National Academy of Sciences of the United States of America, (2020) 117(13), 7290-7295, doi:10.1073/pnas.1915881117

[73] Kobayashi, Y., Wakano, J.Y. & Ohtsuki, H. "Evolution of cumulative culture for niche construction."
Journal of Theoretical Biology, (2019) 472, 67-76, doi:10.1016/j.jtbi.2019.04.013

[72] Uchiumi, Y., Ohtsuki, H. & Sasaki, A. "Evolution of self-limited cell division of symbionts."
Proceedings of the Royal Society B: Biological Sciences, (2019) 286, 20182238, doi:10.1098/rspb.2018.2238

[71] Parvinen, K., Ohtsuki, H. & Wakano, J.Y. "Spatial heterogeneity and evolution of fecundity-affecting traits."
Journal of Theoretical Biology, (2018) 454, 190-204, doi:10.1016/j.jtbi.2018.06.005

[70] Ohtsuki, H. "Evolutionary dynamics of coordinated cooperation."
Frontiers in Ecology and Evolution, (2018) 6, 62, doi:10.3389/fevo.2018.00062

[69] Kobayashi, Y., Wakano, J.Y. & Ohtsuki, H. "Genealogies and ages of cultural traits: An application of the theory of duality to the research on cultural evolution."
Theoretical Population Biology, (2018) 123, 18-27, doi:10.1016/j.tpb.2018.04.007
This paper won the 2020 Marcus W. Feldman Prize for outstanding article published in Theoretical Population Biology, announced at Theoretical Population Biology, (2020) 131, 1, doi.org/10.1016/j.tpb.2019.12.002

[68] Ito, I. M., Ohtsuki, H. & Sasaki, A. "Emergence of opinion leaders in reference networks."
PLoS ONE, (2018) 13(3), e0193983, doi:10.1371/journal.pone.0193983

[67] Nakahashi, W., Ohtsuki, H. "Evolution of emotional contagion in group-living animals."
Journal of Theoretical Biology, (2018) 440, 12-20, doi:10.1016/j.jtbi.2017.12.015

[66] Uchiumi, Y., Ohtsuki, H. & Sasaki, A. "Evolutionary emergence and maintenance of horizontally transmitted mutualism that do not rely on the supply of standing variation in symbiont quality."
Journal of Evolutionary Biology, (2017) 30, 2211-2221, doi:10.1111/jeb.13187

[65] Reeves, T., Ohtsuki, H. & Fukui, S. "Asymmetric public goods game cooperation through pest control."
Journal of Theoretical Biology (2017) 435, 238-247, doi:10.1016/j.jtbi.2017.09.006

[64] Ohtsuki, H. & Innan, H. "Forward and Backward Evolutionary Processes and Allele Frequency Spectrum in a Cancer Cell Population."
Theoretical Population Biology (2017) 117, 43-50, doi:10.1016/j.tpb.2017.08.006

[63] Ohtsuki, H., Wakano, J.Y. & Kobayashi, Y. "Inclusive fitness analysis of cumulative cultural evolution in an island-structured population."
Theoretical Population Biology (2017) 115, 13-23, doi:10.1016/j.tpb.2017.03.001

[62] Sekiguchi, T. & Ohtsuki, H. "Fixation probabilities of strategies for bimatrix games in finite populations."
Dynamic Games and Applications (2017), 7(1), 93-111, doi:10.1007/s13235-015-0170-2

[61] Parvinen, K., Ohtsuki, H. & Wakano, J.Y. "The effect of fecundity derivatives on the condition of evolutionary branching in spatial models."
Journal of Theoretical Biology (2017) 416, 129-143, doi:10.1016/j.jtbi.2016.12.019

[60] Oya, G. & Ohtsuki, H. "Stable polymorphism of cooperators and punishers in a public goods game."
Journal of Theoretical Biology (2017) 419, 243-253, doi:10.1016/j.jtbi.2016.11.012

[59] Inaba, M., Takahashi, N. & Ohtsuki, H. "Robustness of linkage strategy that leads to large-scale cooperation."
Journal of Theoretical Biology (2016) 409, 97-107, doi:10.1016/j.jtbi.2016.08.035

[58] Nakamura, M., Ohtsuki, H. "Optimal decision rules in repeated games where players Infer an opponent's mind via simplified belief calculation."
Games (2016) 7(3), 19, doi:10.3390/g7030019

[57] Tanaka, H., Ohtsuki, H. & Ohtsubo, Y. "The price of being seen to be just: an intention signalling strategy for indirect reciprocity."
Proceedings of the Royal Society B (2016) 283(1835), 20160694, doi:10.1098/rspb.2016.0694

[56] Kobayashi, Y., Ohtsuki, H. & Wakano, J. Y. "Population size vs. social connectedness -- A gene-culture coevolutionary approach to cumulative cultural evolution."
Theoretical Population Biology (2016) 111, 87-95, doi:10.1016/j.tpb.2016.07.001

[55] Kato, S., Ohtsuki, H., Kondo, Y., Nasu, H. & Hongo, H. "Spatial trends of past occupation sites in southwestern part of Kanto region in Jomon Period, Japan -Multivariate analyses of natural environmental factors-"
Theory and Applications of GIS, (2016) 24(1), 23-30.

[54] Morita, M., Ohtsuki, H. & Hiraiwa-Hasegawa, M. "Does Sexual Conflict between Mother and Father Lead to Fertility Decline? A Questionnaire Survey in a Modern Developed Society."
Human Nature, (2016) 27(2), 201-219, doi:10.1007/s12110-016-9254-y

[53] Morita, M., Ohtsuki, H. & Hiraiwa-Hasegawa, M. "A panel data analysis of the probability of childbirth in a Japanese sample: new evidence of the two-child norm."
American Journal of Human Biology, (2016) 28(2), 220-225, doi:10.1002/ajhb.22776

[52] Tonoike, A., Nagasawa, M., Mogi, K., Serpell, J.A., Ohtsuki, H. & Kikusui, T. "Comparison of owner-reported behavioral characteristics among genetically clustered breeds of dog (Canis familiaris)."
Scientific Reports, (2015) 5, 17710, doi:10.1038/srep17710

[51] Ohtsuki, H., Iwasa, Y. & Nowak, M. A. "Reputation effects in public and private interactions."
PLoS Computational Biology, (2015) 11(11), e1004527, doi:10.1371/journal.pcbi.1004527

[50] Sekiguchi, T. & Ohtsuki, H. "Effective group size of majority vote accuracy in sequential decision-making."
Japan Journal of Industrial and Applied Mathematics, (2015) 32(3), 595-614, doi:10.1007/s13160-015-0192-6

[49] Nakahashi, W. & Ohtsuki, H. "When is emotional contagion adaptive?"
Journal of Theoretical Biology (2015) 380, 480-488, doi:10.1016/j.jtbi.2015.06.014

[48] Kobayashi, Y, Wakano, J. Y. & Ohtsuki, H. "A paradox of cumulative culture."
Journal of Theoretical Biology (2015) 379, 79-88, doi:10.1016/j.jtbi.2015.05.002

[47] Ito, K, Ohtsuki, H. & Yamauchi, A. "Relationship between aggregation of rewards and the possibility of polymorphism in continuous snowdrift games."
Journal of Theoretical Biology (2015) 372, 47-53, doi:10.1016/j.jtbi.2015.02.015

[46] So, K. H. Z., Ohtsuki, H. & Kato, T. "Spatial effect on stochastic dynamics of bistable evolutionary games."
Journal of Statistical Mechanics: Theory and Experiment (2014) P10020, doi:10.1088/1742-5468/2014/10/P10020

[45] Nakamura, M. & Ohtsuki, H. "Indirect reciprocity in three types of social dilemmas."
Journal of Theoretical Biology (2014) 355, 117-127, doi:10.1016/j.jtbi.2014.03.035

[44] Ohtsuki, H. "Evolutionary dynamics of n-player games played by relatives."
Philosophical Transactions of the Royal Society B: Biological Sciences (2014) 369(1642), 20130359 doi:10.1098/rstb.2013.0359

[43] Kobayashi, Y. & Ohtsuki, H. "Evolution of social versus individual learning in a subdivided population revisited: Comparative analysis of three coexistence mechanisms using the inclusive-fitness method."
Theoretical Population Biology (2014) 92, 78-87, doi:10.1016/j.tpb.2013.12.003

[42] Mpolya, E. A., Yashima, K., Ohtsuki, H. & Sasaki, A. "Epidemic dynamics of a vector-borne disease on a villages-and-city star network with commuters."
Journal of Theoretical Biology (2014) 343, 120-126, doi:10.1016/j.jtbi.2013.11.024

[41] Úbeda, F. Ohtsuki, H. & Gardner, A. "Ecology drives intragenomic conflict over menopause."
Ecology Letters (2014) 17(2), 165-174, doi:10.1111/ele.12208

[40] Rand, D. G., Tarnita, C. E., Ohtsuki, H. & Nowak, M. A. "Evolution of fairness in the one-shot anonymous Ultimatum Game"
Proceedings of the National Academy of Sciences of the United States of America (2013) 110, 2581-2586, doi:10.1073/pnas.1214167110

[39] Wakano, J. Y., Ohtsuki, H. & Kobayashi, Y. "A mathematical description of the inclusive fitness theory."
Theoretical Population Biology (2013) 84, 46-55, doi:10.1016/j.tpb.2012.11.007

[38] Kashima, K., Ohtsuki, H. & Satake, A. "Fission-fusion bat behavior as a strategy for balancing the conflicting needs of maximizing information accuracy and minimizing infection risk."
Journal of Theoretical Biology (2013) 318, 101-109, doi:10.1016/j.jtbi.2012.10.034

[37] Ohtsuki, H. "Recent developments of evolutionary game theory in finite populations." (Japanese)
Proceedings of the Institute of Statistical Mathematics (2012) 60(2), 251-262

[36] Ohtsuki, H. "Does synergy rescue the evolution of cooperation? -- An analysis for homogeneous populations with non-overlapping generations."
Journal of Theoretical Biology (2012) 307, 20-28, doi:10.1016/j.jtbi.2012.04.030

[35] Morita, M., Ohtsuki, H., Sasaki, A. & Hiraiwa-Hasegawa, M. "Factors affecting the number of children in five developed countries: A statistical analysis with an evolutionary perspective."
Letters on Evolutionary Behavioral Science (2012) 3(1), 7-11, doi:10.5178/lebs.2012.19

[34] Ohtsuki, H. "Evolutionary dynamics of the Nash demand game: a diffusion approach."
Dynamic Games and Applications (2011) 1(3), 449-461, doi:10.1007/s13235-011-0012-9

[33] Cao, L., Ohtsuki, H., Wang, B. & Aihara, K. "Evolution of cooperation on adaptively weighted networks."
Journal of Theoretical Biology (2011) 272(1), 8-15, doi:10.1016/j.jtbi.2010.12.008

[32] Ohtsuki, H. "Evolutionary games in Wright's island model: kin selection meets evolutionary game theory."
Evolution (2010) 64(12), 3344-3353, doi:10.1111/j.1558-5646.2010.01117.x

[31] Bozic, I., Antal, T., Ohtsuki, H., Carter, H., Kim, D., Chen, S., Karchin, R., Kinzler, K. W., Vogelstein, B. & Nowak, M. A. "Accumulation of driver and passenger mutations during tumor progression."
Proceedings of the National Academy of Sciences of the United States of America (2010) 107(43), 18545-18550, doi:10.1073/pnas.1010978107

[30] Rand, D. G., Armao IV, J. J., Nakamaru, M. & Ohtsuki, H. "Anti-social punishment can prevent the co-evolution of punishment and cooperation."
Journal of Theoretical Biology (2010) 265(4), 624-632, doi:10.1016/j.jtbi.2010.06.010

[29] Ohtsuki, H. "Stochastic evolutionary dynamics of bimatrix games."
Journal of Theoretical Biology (2010) 264, 136-142, doi:10.1016/j.jtbi.2010.01.016

[28] Masuda, N. & Ohtsuki, H. "A theoretical analysis of temporal difference learning in the interated Prisoner's Dilemma game."
Bulletin of Mathematical Biology (2009) online first, doi:10.1007/s11538-009-9424-8

[27] Ohtsuki, H. & Nowak, M. A. "Prelife catalysts and replicators."
Proceedings of the Royal Society B (2009) 276, 3783-3790, doi:10.1098/rspb.2009.1136

[26] Tarnita, C. E., Ohtsuki, H., Antal, T., Fu, F. & Nowak, M. A. "Strategy selection in structured populations."
Journal of Theoretical Biology (2009) 259, 570-581, doi:10.1016/j.jtbi.2009.03.035

[25] Tarnita, C. E., Antal, T., Ohtsuki, H. & Nowak, M. A. "Evolutionary dynamics in set structured populations."
Proceedings of the National Academy of Sciences of the United States of America (2009) 106, 8601-8604, doi:10.1073/pnas.0903019106

[24] Antal, T., Ohtsuki, H., Wakeley, J., Taylor, P. & Nowak, M. "Evolution of cooperation by phenotypic similarity."
Proceedings of the National Academy of Sciences of the United States of America (2009) 106, 8597-8600, doi:10.1073/pnas.0902528106

[23] Ohtsuki, H. & Tsuji, K. "Adaptive reproduction schedule as a cause of worker policing in social Hymenoptera: a dynamic game analysis."
The American Naturalist (2009) 173, 747-758, doi:10.1086/598488

[22] Antal, T., Traulsen, A., Ohtsuki, H., Tarnita, C. & Nowak, M. "Mutation-selection equilibrium in games with multiple strategies."
Journal of Theoretical Biology (2009) 258, 614-622, doi:10.1016/j.jtbi.2009.02.010

[21] Rodrigues, A., Koeppl, H., Ohtsuki, H. & Satake, A. "A game theoretical model of deforestation in human-environment relationships."
Journal of Theoretical Biology (2009) 258, 127-134, doi:10.1016/j.jtbi.2009.01.005

[20] Masuda, N. & Ohtsuki, H. "Evolutionary dynamics and fixation probabilities in directed networks."
New Journal of Physics (2009) 11, 033012, doi:10.1088/1367-2630/11/3/033012

[19] Manapat, M., Ohtsuki, H., Bürger, R. & Nowak, M. A. "Originator dynamics."
Journal of Theoretical Biology (2009) 256, 586-595, doi:10.1016/j.jtbi.2008.10.006

[18] Rand, D., Ohtsuki, H. & Nowak, M. "Direct reciprocity with costly punishment: generous tit-for-tat prevails."
Journal of Theoretical Biology (2009) 256, 45-57, doi:10.1016/j.jtbi.2008.09.015

[17] Ohtsuki, H., Iwasa, Y. & Nowak, M. A. "Indirect reciprocity provides only a narrow margin of efficiency for costly punishment."
Nature (2009) 457, 79-82, doi:10.1038/nature07601

[16] Nowak, M. A. & Ohtsuki, H. "Prevolutionary dynamics and the origin of evolution."
Proceedings of the National Academy of Sciences of the United States of America (2008) 105, 14924-14927, doi:10.1073/pnas.0806714105

[15] Ohtsuki, H. & Nowak, M. A. "Evolutionary stability on graphs."
Journal of Theoretical Biology (2008) 251, 698-707, doi:10.1016/j.jtbi.2008.01.005

[14] Pacheco, J. M., Traulsen, A., Ohtsuki, H. & Nowak, M. A. "Repeated games and direct reciprocity under active linking."
Journal of Theoretical Biology (2008) 250, 723-731, doi:10.1016/j.jtbi.2007.10.040

[13] Ohtsuki, H.*, Bordalo, P.* & Nowak, M. A. (* joint first-authors) "The one-third law of evolutionary dynamics."
Journal of Theoretical Biology (2007) 249, 289-295, doi:10.1016/j.jtbi.2007.07.005

[12] Ohtsuki, H. & Nowak, M. A. "Direct reciprocity on graphs."
Journal of Theoretical Biology (2007) 247, 462-470, doi:10.1016/j.jtbi.2007.03.018

[11] Ohtsuki, H., Pacheco, J. & Nowak, M. A. "Evolutionary graph theory: Breaking the symmetry between interaction and replacement."
Journal of Theoretical Biology (2007) 246, 681-694, doi:10.1016/j.jtbi.2007.01.024

[10] Uehara, T., Iwasa, Y. & Ohtsuki, H. "ESS distribution of display duration in animal contests to assess an opponent before fighting or fleeing."
Evolutionary Ecology Research (2007) 9, 395-408

[9] Ohtsuki, H., Nowak, M. A. & Pacheco, J. "Breaking the symmetry between interaction and replacement in evolutionary dynamics on graphs."
Physical Review Letters (2007) 98, 108106, doi:10.1103/PhysRevLett.98.108106
(also appeared in Virtual Journal of Biological Physics Research (2007) 13(6))

[8] Masuda, N. & Ohtsuki, H. "Tag-based indirect reciprocity by incomplete social information."
Proceedings of the Royal Society B: Biological Sciences (2007) 274, 689-695, doi:10.1098/rspb.2006.3759

[7] Ohtsuki, H. & Iwasa, Y. "Global analyses of evolutionary dynamics and exhaustive search for social norms that maintain cooperation by reputation."
Journal of Theoretical Biology (2007) 244, 518-531, doi:10.1016/j.jtbi.2006.08.018

[6] Ohtsuki, H. & Nowak, M. A. "The replicator equation on graphs."
Journal of Theoretical Biology (2006) 243, 86-97, doi:10.1016/j.jtbi.2006.06.004

[5] Ohtsuki, H. & Nowak, M. A. "Evolutionary games on cycles."
Proceedings of the Royal Society B: Biological Sciences (2006) 273, 2249-2256, doi:10.1098/rspb.2006.3576

[4] Ohtsuki, H., Hauert, C., Lieberman, E. & Nowak, M. A. "A simple rule for evolution of cooperation on graphs and social networks."
Nature (2006) 441, 502-505, doi:10.1038/nature04605

[3] Ohtsuki, H. & Iwasa, Y. "The leading eight: social norms that can maintain cooperation by indirect reciprocity."
Journal of Theoretical Biology (2006) 239, 435-444, doi:10.1016/j.jtbi.2005.08.008

[2] Ohtsuki, H. & Iwasa, Y. "How should we define goodness? - reputation dynamics in indirect reciprocity."
Journal of Theoretical Biology (2004) 231, 107-120, doi:10.1016/j.jtbi.2004.06.005
[Erratum] Ohtsuki, H. & Iwasa, Y. "Erratum to: How should we define goodness?-reputation dynamics in indirect reciprocity [Journal of Theoretical Biology 231 (2004) 107-120]" Journal of Theoretical Biology (2005) 232(4), 451, doi:10.1016/j.jtbi.2004.10.001

[1] Ohtsuki, H. "Reactive strategies in indirect reciprocity."
Journal of Theoretical Biology (2004) 227, 299-314, doi:10.16/j.jtbi.2003.11.008
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